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The Philippine Agricultural Scientist

Rommel C. Sulabo, Ph.D. 
Editor-in-Chief 
Institute of Animal Science 
College of Agriculture and Food Science 

Barbara L. Caoili, Ph.D. 
Executive Editor 
National Crop Protection Center 
College of Agriculture and Food Science 

Merdelyn C. Lit, Ph.D. 
Technical Editor 
Institute of Plant Breeding 
College of Agriculture and Food Science 

Lourdes B. Columbres, MBA 
Associate Editor 

Anacorita O. Abasolo, M.Sc. 
Production Editor 

Maria Dominique L. Culing 
Language Editor 

Clarissa Q. Belen 
Administrative Support Staff 

Section Editors 

Barbara L. Caoili, Ph.D. 
National Crop Protection Center 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Merdelyn C. Lit, Ph.D. 
Institute of Plant Breeding 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Jose E. Hernandez, Ph.D. 
Institute of Crop Science 
College of Agriculture and Food Science, 
University of the Philippines Los Baños 

Pompe C. Sta Cruz, Ph.D. 
Institute of Crop Science 
College of Agriculture and Food Science, 
University of the Philippines Los Baños 

Virginia R. Cardenas, Ph.D. 
Institute of Governance and Rural Development 
College of Public Affairs and Development 
University of the Philippines Los Baños 

Jessamyn R. Adorada, Ph.D. 
Institute of Weed Science, Entomology and Plant Pathology 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Mudjekeewis D. Santos, Ph.D. 
Academician, 
National Fisheries Research and Development Institute 
Diliman, Quezon City 

Kevin F. Yaptengco, Ph.D. 
College of Engineering and Agro-Industrial Technology, 
University of the Philippines Los Baños 

Filomena C. Sta cruz, Ph.D. 
Institute of Weed Science, Entomology and Plant Pathology 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Elda B. Esguerra, Ph.D. 
Postharvest Horticulture Training and Research Center 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Floirendo P. Flores, Ph.D. 
Institute of Food Science and Technology 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Consuelo Amor S. Estrella, Ph.D. 
Institute of Animal Science 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Analiza Henedina M. Ramirez, Ph.D. 
Institute of Weed Science, Entomology and Plant Pathology 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Crisanta S. Bueno, Ph.D. 
Institute of Crop Science 
College of Agriculture and Food Science 
University of the Philippines Los Baños 

Tonette P. Laude, Ph.D. 
Institute of Crop Science 
College of Agriculture and Food Science, 
University of the Philippines Los Baños 

Consorcia E. Reaño, Ph.D. 
Institute of Crop Science 
College of Agriculture and Food Science, 
University of the Philippines Los Baños 

Roger A. Luyun, Ph.D. 
College of Engineering and Agro-Industrial Technology, 
University of the Philippines Los Baños 

Rubenito M. Lampayan, Ph.D. 
College of Engineering and Agro-Industrial Technology, 
University of the Philippines Los Baños 

Ma. Cristine Concepcion D. Ignacio, Ph.D. 
College of Engineering and Agro-Industrial Technology, 
University of the Philippines Los Baños 

Percival P. Sangel, Ph.D. 
Institute of Animal Science, 
College of Agriculture and Food Science, 
University of the Philippines Los Baños 

Maria Ana T. Quimbo, Ph.D. 
Institute of Governance and Rural Development, 
College of Public Affairs and Development, 
University of the Philippines Los Baños 

Dinah Pura T. Depositario, Ph.D. 
Department of Agribusiness Management and Entrepreneurship, 
College of Economics and Management, 
University of the Philippines Los Baños 

Editorial Board 

Jose Ramon Botella, Ph.D. 
School of Biological Sciences 
University of Queensland 
Brisbane, Queensland, Australia 

Ian Coxhead, Ph.D. 
Department of Agricultural and Applied Ecoomics 
University of Wisconsin 
Madison, Wisconsin, USA 

Fabian M. Dayrit, Ph.D. 
College of Science 
Ateneo de Manila University 
Quezon City, Philippines 

Ernesto J. Del Rosario, Ph.D. 
(Professor Emeritus) Institute of Chemistry 
University of the Philippines Los Baños 
College Laguna, Philippines 

Virgilio A. Fernandez, Ph.D. 
College of Forestry 
University of the Philippines Los Baños 
College Laguna, Philippines 

Jane Haggis, Ph.D. 
Center for Development Studies 
Flinders University 
Adelaide, South Australia 

Editorial Consultative Board 

Arnel N. Del Barrio, Ph.D. 
 

Dolores A. Ramirez, Ph.D. 
 

Rafael D. Guerrero, Ph.D. 
 

Evelyn Mae Tecson-Mendoza, Ph.D. 
 

National Academy of Science and Technology 
Department of Science and Technology (DOST) 
Bicutan Tguig City, Metro Manila Philippines 
email: secretariat@nast.ph 

Teresita L. Rosario, Ph.D. 
 

Ofelia K. Bautista, Ph.D. 
 

Calixto M. Protacio, Ph.D. 
 

Constancio C. De Guzman, Ph.D. 
 

Christian Joseph R. Cumagun, Ph.D.