The Philippine Agricultural Scientist
Rommel C. Sulabo, Ph.D.
Editor-in-Chief
Institute of Animal Science
College of Agriculture and Food Science
Barbara L. Caoili, Ph.D.
Executive Editor
National Crop Protection Center
College of Agriculture and Food Science
Merdelyn C. Lit, Ph.D.
Technical Editor
Institute of Plant Breeding
College of Agriculture and Food Science
Lourdes B. Columbres, MBA
Managing Editor
Anacorita O. Abasolo, M.Sc.
Production Editor
Maria Dominique L. Culing
Language Editor
Clarissa Q. Belen
Administrative Support Staff
Section Editors
Barbara L. Caoili, Ph.D.
Executive Editor
National Crop Protection Center
College of Agriculture and Food Science
Merdelyn C. Lit, Ph.D.
Technical Editor
Institute of Plant Breeding
College of Agriculture and Food Science
Jose E. Hernandez, Ph.D.
Institute of Crop Science
College of Agriculture and Food Science,
University of the Philippines Los Baños
Pompe C. Sta Cruz, Ph.D.
Institute of Crop Science
College of Agriculture and Food Science,
University of the Philippines Los Baños
Virginia R. Cardenas, Ph.D.
Institute of Governance and Rural Development
College of Public Affairs and Development
University of the Philippines Los Baños
Jessamyn R. Adorada, Ph.D.
Institute of Weed Science, Entomology and Plant Pathology
College of Agriculture and Food Science
University of the Philippines Los Baños
Mudjekeewis D. Santos, Ph.D.
Academician,
National Fisheries Research and Development Institute
Diliman, Quezon City
Kevin F. Yaptengco, Ph.D.
Department of Agricultural Bio Processing
College of Engineering and Agro-Industrial Technology,
University of the Philippines Los Baños
Filomena C. Sta cruz, Ph.D.
Institute of Weed Science, Entomology and Plant Pathology
College of Agriculture and Food Science
University of the Philippines Los Baños
Elda B. Esguerra, Ph.D.
Postharvest Horticulture Training and Research Center
College of Agriculture and Food Science
University of the Philippines Los Baños
Floirendo P. Flores, Ph.D.
Institute of Food Science and Technology
College of Agriculture and Food Science
University of the Philippines Los Baños
Consuelo Amor S. Estrella, Ph.D.
Institute of Animal Science
College of Agriculture and Food Science
University of the Philippines Los Baños
Analiza Henedina M. Ramirez, Ph.D.
Institute of Weed Science, Entomology and Plant Pathology
College of Agriculture and Food Science
University of the Philippines Los Baños
Honorary Advisory Board
Clare R. Baltazar, Ph.D.
Academician, National Academy of Science and Technology
Philippine Department of Science and Technology
Evelyn Mae Tecson-Mendoza, Ph.D.
Academician, National Academy of Science and Technology
Philippine Department of Science and Technology
Ricardo M. Lantican, Ph.D.
Academician, National Scientists
Alumni of the College of Agriculture
University of the Philippines Los Baños
Rafael D. Guerrero, Ph.D.
Academician, National Academy of Science and Technology
Philippines Department of Science and Technology (DOST)
Bicutan, Taguig City, Metro Manila,
Philippines; e-mail: secretariat@nast.ph
Arnel N. Del Barrio, Ph.D.
Academician, National Academy of Science and Technology
University of the Philippines Los Baños
Editorial Consultative Board
Jose Ramon Botella, Ph.D.
School of Biological Sciences,
University of Queensland Brisbane,
Queensland, Australia
Ian Coxhead, Ph.D.
Department of Agricultural and Applied Economics
University of Wisconsin
Madison, Wisconsin, U. S. A.
Fabian M. Dayrit, Ph.D.
College of Science,
Ateneo de Manila University
Quezon City, Philippines
Salcedo L. Eduardo, Ph.D.
College of Veterinary Medicine
University of the Philippines Los Baños
Jane Haggis, Ph.D.
Center for Development Studies,
Flinders University Adelaide, South Australia
Ernesto J. Del Rosario, Ph.D. (Professor Emeritus)
Institute of Chemistry
University of the Philippines Los Baños